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Charpentiera tomentosa

Sohmer (1972)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-CoreEudicot Order: Caryophyllales Family: Amaranthaceae Genus: Charpentiera

pāpala [papala]

Description

Key Characters:

Growth Form: Slender or robust trees 6–12 m tall, new growth densely brown tomentose.

Stems: Stems 1.75–6 mm in diameter directly below apex, internodes up to 12 mm long.

Roots:

Leaves: Leaves simple. Alternate. Blades obovate, elliptic, or subovate, (8.5–)15–30 cm long, 2.6–12 cm wide. Apex acute to rounded and usually retuse as well. Base acute to rounded. Surfaces glabrous or pubescent and stiffly coriaceous. Margins entire. Lateral veins 10–20 pairs. Marginal ribs strongly and prominently developed. Petioles 1–11 cm long. Stipules absent.

Flowers: Flowers in single axillary panicles, these 1–2 branched, reddish brown pubescent or glabrous. Panicles 13–50 cm long, often compound, with 8–25 primary branches 5–20 cm long. Each panicle branch subtended by 1 bract, peduncles 2–10 cm long. Gynodioecious (with bisexual (perfect)–flowered plants and female plants in each population). flowers several to 20–30 per branch, internodes between individual flowers 0.8–11 mm long. Calyx of 5 sepals, distinct, imbricate, subequal; 1.5–2.5 mm long, 3–5-veined. Corolla (petals) absent. Stamens and staminodes 5; filaments connate at base to form a short, membranous tube with short, rounded flaps of tissue alternating with the filaments; anthers exserted at anthesis. Ovary superior, globose or urceolate, 1.5–2 mm long; style absent or very short; stigmas 2, straight or recurved, nearly as long as the ovary, with glands on the inner surface.

Fruit: Indehiscent utricle. Seeds black and shiny; lenticular ca. 1.5 mm long.

Ploidy:

Habitat: Occuring in mesic forest.

Elevation Range: 110–770 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

O'ahu Endemic
Molokai Endemic
Lana'i Endemic
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: Brittonia 24: 300 (1972)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Charpentiera tomentosa Sohmer upper makaha valley, waianae range, off crest of makaha-waianae kai ridge on the makaha side, beyond the trail coming up from makaha valley aa lava, jungle PRESERVED_SPECIMEN obata, j.k. palmer, d.; funk, e.; hall, j. collector number: 339 Hawaii BISH 2/19/1978
2 Charpentiera tomentosa Sohmer waianae mts., mokuleia-makua divide, mokulera forest bern Mokuleia rainforest at PRESERVED_SPECIMEN st.john, h. sohmer, s.h. collector number: 12241 Oahu BISH 11/25/1932
3 Charpentiera tomentosa Sohmer pahole gulch; drainage #2 Rare vine climbing 10' into Diospyros in old aa flow with Diospyros-Canthium-Lantana-Pennisetum association. PRESERVED_SPECIMEN welton, p. collector number: 102 Hawaii BISH 3/17/1989
4 Charpentiera tomentosa Sohmer Waianae Kai Trail Diverse mesic forest. PRESERVED_SPECIMEN Annable, C.R. Ray, G.J. Collector Number: 3864 Oahu BISH 10/2/1998
5 Charpentiera tomentosa Sohmer puu kanehoa, honouliuli, waianae mts Wooded ridge PRESERVED_SPECIMEN st.john, h. collector number: 11055 Oahu BISH 3/22/1931
6 Charpentiera tomentosa Sohmer kawainui ditch trail On shaded hillside, PRESERVED_SPECIMEN crosby, m.r. anderson, w.r. collector number: 1572 Oahu BISH 6/28/1964
7 Charpentiera tomentosa Sohmer kaiholena PRESERVED_SPECIMEN munro, g.c. collector number: 44 Lanai BISH 8/8/1913
8 Charpentiera tomentosa Sohmer waianae mts Sparse Styphalia, Dodonaea, Vaccinium vegatation. PRESERVED_SPECIMEN long, c.r. herbst, d.r. collector number: 4016 long Hawaii BISH 1/18/1966
9 Charpentiera tomentosa Sohmer maakua gulch PRESERVED_SPECIMEN porter, j. collector number: 3 Oahu BISH 10/18/1969
10 Charpentiera tomentosa Sohmer makaha valley, waianae mts Lower woods PRESERVED_SPECIMEN st.john, h. collector number: 11619 Oahu BISH 4/3/1932
11 Charpentiera tomentosa Sohmer northern waianae mts., peacock flats trail; trail begins about Winds through forest with Koa, the native Hibiscus, Pisonia, and Charpentiera among others. PRESERVED_SPECIMEN sohmer, s.h. collector number: 6653 Oahu BISH 9/5/1970
12 Charpentiera tomentosa Sohmer waianae kai trail Diverse mesic forest. PRESERVED_SPECIMEN annable, c.r. ray, g.j. collector number: 3864 Oahu BISH 10/2/1998
13 Charpentiera tomentosa Sohmer waianae mts., mokuleia-makua divide, mokulera forest bern PRESERVED_SPECIMEN st.john, h. sohmer, s.h. collector number: 12241 Oahu BISH 11/25/1932
14 Charpentiera tomentosa Sohmer upper eaches of manini gulch Gulch PRESERVED_SPECIMEN takeuchi, w.n. pyle, l.; tate, b. collector number: 2311 Oahu BISH 7/14/1985
15 Charpentiera tomentosa Sohmer waianae saddle between puu kawiwi and mt. kaala PRESERVED_SPECIMEN montgomery, s.l. collector number: s.n. Oahu BISH 6/5/1970
16 Charpentiera tomentosa Sohmer waianae valley, gulches west of aianae kai below puu kawiwi PRESERVED_SPECIMEN perlman, s.p. obata, j. collector number: 5501 Maui BISH 3/8/1987
17 Charpentiera tomentosa Sohmer kaliuwaa PRESERVED_SPECIMEN takeuchi, w.n. collector number: 3916 Oahu BISH 7/30/1988
18 Charpentiera tomentosa Sohmer koolau mts., hidden valley above kaaawa - lower part of valley Kukui, Awa patches, Banana on stream bank by waterfall, south side of stream PRESERVED_SPECIMEN perlman, s.p. obata, j. collector number: 5413 Oahu BISH 3/8/1987
19 Charpentiera tomentosa Sohmer kawaiiki, typical kukui-clidemia gully Distributes preferentially along the bottoms and lower slopes of area gullies PRESERVED_SPECIMEN takeuchi, w.n. paquin collector number: 3030 Oahu BISH 11/22/1986
20 Charpentiera tomentosa Sohmer puu kalena, waianae mts. waianae kai Wooded gulch PRESERVED_SPECIMEN st.john, h. collector number: 12975 Oahu BISH 3/12/1933
21 Charpentiera tomentosa Sohmer kaluanui PRESERVED_SPECIMEN takeuchi, w.n. paquin collector number: 39321 Kauai US 7/31/1988
22 Charpentiera tomentosa Sohmer northern waianae mts., peacock flats trail; trail begins about In scrub, on trailside Metrosideros. PRESERVED_SPECIMEN sohmer, s.h. collector number: 6652 Maui BISH 9/5/1970
23 Charpentiera tomentosa Sohmer oahu, ka uaa gulch, waianae mountains PRESERVED_SPECIMEN s. h. sohmer et al. 6438 MICH 5/27/1970
24 Charpentiera tomentosa Sohmer waianae kai trail Diverse mesic forest PRESERVED_SPECIMEN c. r. annable 3864 NY 10/2/1998
25 Charpentiera tomentosa Sohmer oahu: waimea arboretum. Cultivated garden PRESERVED_SPECIMEN james s. miller|mary merello miller 7879 NY 7/31/1992
26 Charpentiera tomentosa Sohmer island of oahu. waianae kai trail, waianae kai forest reserve, west fork of western-most tributary of kaupuni stream, along stream banks and up slopes Diverse mesic forest with aliens invading PRESERVED_SPECIMEN g. j. ray 1047 NY 10/2/1998
27 Charpentiera tomentosa Sohmer Waianae Mts, Waianae Valley, gulches west of Waianae K ai below Puu Kawiwi. In Sapindus forest. PRESERVED_SPECIMEN Steve Perlman 5501 Oahu PTBG 8/3/1987
28 Charpentiera tomentosa Sohmer Waianae Mts; Kahanahaiki Army Ecosystem Management Region; Makua. Diospyros hillebrandii lowland mesic forest. PRESERVED_SPECIMEN K. R. Wood 7714-A Oahu PTBG 3/11/1999
29 Charpentiera tomentosa Sohmer Waianae Mts; Kahanahaiki Army Ecosystem Management Region; Makua. Elev. 1820 ft. Diospyros hillebrandii lowland mesic forest. PRESERVED_SPECIMEN K. R. Wood 7714-B Oahu PTBG 3/11/1999
30 Charpentiera tomentosa Sohmer Waianae Mountains, Kaluaa Gulch. PRESERVED_SPECIMEN Sohmer, S. H. 6438 Oahu US 5/27/1970
31 Charpentiera tomentosa Sohmer Oahu: Waimea Arboretum. PRESERVED_SPECIMEN James S. Miller|Mary Merello Miller 7879 Missouri Botanical Garden 7/31/1992
32 Charpentiera tomentosa Sohmer Waianae Kai Trail Diverse mesic forest PRESERVED_SPECIMEN C. R. Annable 3864 NY 10/2/1998
33 Charpentiera tomentosa Sohmer island of oahu. waianae kai trail, waianae kai forest reserve, west fork of western-most tributary of kaupuni stream, along stream banks and up slopes PRESERVED_SPECIMEN g. j. ray 1047 NY 10/2/1998
34 Charpentiera tomentosa Sohmer Koolau Mts, Hidden Valley above Kaaawa, in lower part of Hidden Valley near awa patches, banana, taro on stream bank near waterfall near Schiedea kaalae #5414 south side of stream. PRESERVED_SPECIMEN Steve Perlman 5413 Oahu PTBG 8/3/1987
35 Charpentiera tomentosa Sohmer wailele Emerging from water seep along vertical basalt face PRESERVED_SPECIMEN takeuchi, w.n. yoshinaga, a. collector number: 2939 Oahu BISH 10/1/1986
36 Charpentiera tomentosa Sohmer Honolulu Harbor PRESERVED_SPECIMEN Sohmer, S.H. Collector Number: 6683 Oahu BISH 1/1/1970
37 Charpentiera tomentosa Sohmer honolulu harbor PRESERVED_SPECIMEN sohmer, s.h. collector number: 6683 Oahu BISH 8/8/1970
38 Charpentiera tomentosa Sohmer kaliuwaa Narrow flood plain with monodominant forest of Syzygium malaccense, loamy substrate infused with alluvial pebbles along 1% streamcourse PRESERVED_SPECIMEN takeuchi, w.n. collector number: 3916 Oahu BISH 7/30/1988
39 Charpentiera tomentosa Sohmer wiliwilinui ridge, e slope PRESERVED_SPECIMEN tokushige, e. collector number: s.n. Oahu BISH 6/28/1970
40 Charpentiera tomentosa Sohmer makaha valley, waianae range, off crest of makaha-waianae kai ridge on the makaha side beyond the old trail coming up from makaha valley; under a canopy of natives: metrosideros, euginia, etc subalpine shrubland, SW facing hillside in weathered aa with sandy soil layer; Sophora PRESERVED_SPECIMEN obata, j.k. palmer, j.; funk, e.; hall, j. collector number: 339 Hawaii BISH 2/19/1978
41 Charpentiera tomentosa Sohmer maakua gulch PRESERVED_SPECIMEN porter, j. collector number: 2 Hawaii BISH 10/18/1969
42 Charpentiera tomentosa Sohmer head of Hulopoe Gulch at Munro trail PRESERVED_SPECIMEN Degener, O.; I.Degener BISH 8/11/1963
43 Charpentiera tomentosa Sohmer waianae mts., along access rd. to kuaokala forest reserve, head of kawaihapi gulch Gulch - Steep slope dominated by {Pisonia and Charpentiera PRESERVED_SPECIMEN wagner, w.l. takeuchi, w.; obata, j. collector number: 5355 Oahu BISH 4/8/1984
44 Charpentiera tomentosa Sohmer waianae mts., mokuleia forest reserve, pahole gulch, 1st eastern subgulch Steep rocky slope in mixed mesophytic forest with Pisonia, Hedyotis, Gouldia, Psychotria, Charpentiera, Morinda, and Hibiscus PRESERVED_SPECIMEN wagner, w.l. takeuchi, w.; obata, j. collector number: 5345 Oahu BISH 4/8/1984
45 Charpentiera tomentosa Sohmer waianae kai, gulch at base of kamaileunu ridge Gulch in guava-kukui forest PRESERVED_SPECIMEN takeuchi, w.n. collector number: 2109 Oahu BISH 3/31/1985
46 Charpentiera tomentosa Sohmer kaluanui Syzygium malaccense forest along Cyrtandra-rich streamcourse, on clay banks or talus rock PRESERVED_SPECIMEN takeuchi, w.n. paquin collector number: 39321 Oahu BISH 7/31/1988
47 Charpentiera tomentosa Sohmer kawaiiki ditch trail, koolau mountains PRESERVED_SPECIMEN crosby, m.r. anderson, w.r. collector number: 1941 Oahu BISH 8/1/1964
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